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GR can perform its enzymatic activity in the cytoplasm, but also in the ER and within lysosomes, mitochondria and the nucleus

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doi:10.1158/1535-7163.MCT-05-0523 Troussard AA, Costello P, Yoganathan EN, Kumagai S, Roskelley CD, Dedhar S (2000) The integrin linked kinase (ILK) induces an invasive phenotype via AP-1 transcription factor-dependent upregulation of matrix metalloproteinase 9 (MMP-9)