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These contrasting responses indicate that flavonoid reprogramming under combined stress depends not only on stress composition but also on species-specific regulatory architectures and metabolic capacities, limiting the transferability of engineering strategies across species ( Combined stress additionally reshapes primary carbon metabolism, thereby influencing substrate availability for phenylpropanoid biosynthesis

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It undergoes significant first-pass metabolism in the liver and intestines, where the enzyme arginase breaks it down, reducing bioavailability